Alexander Dennis Enviro500 Open Top North America '2005–11

Gray Line New York, owned by Coach USA ordered 20 open-top Enviro500 for its sightseeing operations in New York and put them into service in spring 2005, these were the first open-top Enviro500 built (another 13 would later be ordered). Gray Line also received three single-door Enviro500 in summer 2005 for operation in San Francisco.

Citizens Area Transit (now RTC Transit) of Las Vegas had its first 50 Enviro500s enter service by October 2005. The service, marketed as "The Deuce", runs along the Las Vegas Strip. Certain high-traffic routes, such as along Maryland Parkway and Nellis Boulevard, have also used these buses. They offer extra ventilation grilles outside the engine compartment to cope with Southern Nevada's dry desert climate and high summertime temperatures. Las Vegas's Enviro500s have the largest air conditioning system ever created for a transit vehicle, with eight fans fitted on the air conditioning unit. As of October 2006, CAT had firm orders for another 41 of the type worth $29.8 million; these were delivered in late 2007. A further 40 had also been ordered for delivery in 2008; these were 12.8 meters long and had a second staircase to speed loading and unloading. All these vehicles were used for regular fixed-route operations, not for sightseeing operations.

Les Cars Rouges, a sightseeing company, ordered 16 open-top Enviro500 for sightseeing operations in the USA, delivery of these buses started in early 2007. Eight of the Enviro500 are destined for San Francisco, and the rest of them are destined for Washington D.C.

Community Transit currently has a fleet of 23 12.8m buses, all of them were introduced in 2011.

Unitrans (the student-run transit service of University of California, Davis, known for years for its operation of former London Transport double-decker buses) ordered two Enviro500 which were delivered in early 2010; these are the first batch of Enviro500s with the bodywork assembled by ElDorado National.

SLO Transit ordered one Enviro500 which entered service in late 2010.

Hele-On Bus introduced one 12.8m Enviro500 in the first half of 2011.

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